Skull squadron is one of two GDI military units and taking charge of its aircraft. Four of the A-15 Orcas were from this squadron as the other four belonged to the Ghost squadron. Their headquarters is located at an Airfield where Nod forces stood their hold and started attacking the two Airfields at Langley Air Force Base. When GDI reclaimed the Airfields, the squadron reunited at this base and attacked Nod's primary garrison by destroying key structures prior to the final objective. Orcas were now deployed at Alexandria in Egypt where they suddenly destroyed two Battleships and loading cranes which are in the hands of Nod with three compounds for controlling the city's region in a dangerous Red Zone.

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